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Vanitas and Noé Archiviste head out to the town of Gévaudan in search of the "Beast," an enormous wolf-like creature that has slaughtered hundreds of people. Suspecting that the Beast is a curse-bearing vampire, Vanitas primarily aims to heal it using the powers of his grimoire. Along the way, the two get separated and suddenly travel back to the past—to the exact moment the Beast is lurking in the woods. After a battle against the gigantic wolf and a vampire hunter, Vanitas decides to team up with Jeanne in order to find Noé. Despite being allies, Jeanne's goal is the opposite of Vanitas', as she was tasked to kill the Beast—suspecting it may be someone she used to know. Meanwhile, a severely wounded Noé is picked up by the mysterious Chloé d'Apchier and her servant. Like Noé, Chloé is a vampire whose existence was erased from the public's knowledge. She has been a guardian for future generations and once tried to find a way to become human again. While Noé is grateful to Chloé for her hospitality, little does he know that she might be siding with forces far more dangerous than the Beast itself. [Written by MAL Rewrite]

Despite their differences in position, three men—the youngest senior executive of the Port Mafia, Osamu Dazai; the lowest ranking member, Sakunosuke Oda; and the intelligence agent, Angou Sakaguchi—gather at the Lupin Bar at the end of the day to relax and take delight in the company of friends. However, one night, Ango disappears. A photograph taken at the bar is all that is left of the three together. Fast forward to the present, and Dazai is now a member of the Armed Detective Agency. The Guild, an American gifted organization, has entered the fray and is intent on taking the Agency's work permit. They must now divide their attention between the two groups, the Guild and the Port Mafia, who oppose their very existence. [Written by MAL Rewrite]