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Underneath the surface of the Earth lies the Inner World, a realm where a war rages on between intelligent spiders and the invading humanoid insects known as Insectors. However, the spiders are not alone, as "Spider Riders"—humans from the Outer World, known as Earthens in the Inner World, that are chosen by the Oracle—fight alongside them. Filled with wondrous tales of a mysterious pyramid and entrusted with his grandfather's journal, a young boy called Hunter Steele sets out on an exploration. There, his destined meeting with the Oracle occurs, and he is transported into the Inner World as a Spider Rider. Finding himself lost and confused in the middle of a forest, Hunter crosses paths with a giant spider named Shadow. Although reluctant, Shadow agrees to fight by his side. He then meets Corona—a skilled Spider Hunter and kind girl with a penchant for normal life. She shows him the ropes of being a Spider Rider, and educates him about the fate of the Inner World if the Insectors continue their tyrannical reign. Thus begins the story of Hunter Steele, a heedless young boy on a quest to save the Inner World. But his task of retrieving the four Oracle Keys holding the Oracle's true power will lead him to an ultimate clash against Mantid, leader of the Insectors. [Written by MAL Rewrite]

Pushed out to the far reaches of the galaxy, the crew of the Solo Ship faces impossible odds of survival. With the Earth destroyed and the remnants of the Buff Clan pushing them to their limit, the Ideon shows as the last line of defense for the battered remains of humanity. As the tensions of their last stand rack up on their minds, Captain Bes Jordan and pilot Cosmo Yuuki race against the clock to protect their final opportunity of survival, even as desperation eats away at the minds of their loved ones. Using any weapons left at their disposal, the crew desperately fight on to protect themselves in the cold clutches of space, punished with a final duel against Harulu Ajiba, who is now desperate to reclaim the honor lost in her father's eyes. [Written by MAL Rewrite]

In a distant future there is an organization called Deava which had to fight against the so called "Shadow Angels", creatures that had survived for over 12,000 years whose only purpose was to destroy humanity. Apollo, a seemingly normal human, was in fact the reincarnation of a fallen Angel called Apollonius, who in turn had to fight against Touma, a Shadow Angel, along with his comrades in order to prevent the destruction of the Earth. (Source: ANN)

Sousei no Aquarion Evol is set in the spring of 1966. Shin Tsukishima, who is travelling to get a crystal radio, meets Yuno Kawazu, a girl crying on a beach in Izu. Shin and Yuno are guided by the Book of Genesis and meet each other halfway. The boundaries between three worlds begin to distort. (Source: ANN) Original episode that will be screened at Project Aquarion's 10th Anniversary event scheduled to be held on June 28, 2015.